When Toland speaks of the three Queens I believe he is speaking of oryx and his sister's and their individual approaches to sword logic. And when he speaks of the single queen living in harmony it is my belief he is referring to us, the traveler and the guardians. And although Toland doesn't believe us capable of defeating the hive and ultimately overcoming death I disagree. We will through cooperation achieve what no single entity could hope to. The deep seeks the final form while the sky rejects the ending altogether.